Barry's web page for host-parasite metapopulation research project


Equations used within this model

Verbal descriptions of the analytic results, figures from corresponding simulations included.


Parameter intensity and bifurcation diagrams A small collection of bifurcation plots (cis-configuration) showing the effects selection coefficients iterating 1 or both migration rates.

Preliminary Bifurcation Diagrams This was the first attempt at a bifurcation diagram of the model. It is a simple 2-island system holding parasite migration at 10%. Shows a zoom in of the point where the polymorphic equilibria is believed to become unstable, a simple bifurcation pattern is not believed to exist.


Parameter intensity as a linear function on period Some plots showing (qualitatively) that halving all alphas and betas appear to halve the periodicity of the system. Need to look at linearized model and see how accurate it is at predicting the period.


2 Island Stability Gallery A collection of stability plots for a 2-island system with 81 plots (each containing ~1500 points) showing stability or lack of stability at the stable polymorphic equilibrium. Some spiffy preliminary observations within.

 

Another 2 island stability gallery Some more 2-island stability plots but with magnitude of the largest eigenvalue instead of just stable or unstable.

 

A 4-island stability gallery Some 4-island stability plots. Interesting to note that that the plots are still symmetric with respect to parasite and host migration rates. Also note that the range of the stable polymorphic equilibrium can be discontinuous.


Initial starting frequencies do not appear to affect long-term behavior

Initial frequencies. Emerging limit cycle or equilibrium is independent of initial allele frequencies, suggesting only 1 behavior in the system is stable.


Observations on Local Adaptation
 
       If the island system exhibits cycling- A species on an island with a strong selection coefficient will be locally adapted. This appears
                 to be from the species "leading" the oscillations and switching before other islands. These islands still often experience
                 periods of local maladaptation.

       If the island is at equilibrium- A species has a very simple expression for local adaptation. A species on an island with a weak
                 selection coefficient will be locally adapted. Further the local adaptation across all islands will sum to zero for each species.

       Local adaptation of the entire island system across time even when cycles are present is extremely low (below 0.1% in almost all
                 cases). Some island systems produce moments of an adapted system but this is usually followed
                 quickly by a period of maladaptation.


Gallery

This is a gallery for a collection of phase plots with a combined frequency over time plot.

 Gallery moved here.



Alternate LA This was an investigation to see if weighting local adaptation by the product of the allele frequencies produced a significant change in local adaptaion levels within the system. It reduced local adaptation by around 6-fold on average, never increasing local adaptation values. The weighted local adaptation values are marked with an extra "*".

LA observations this has the analyticly derived local adaptation values for a  system at equilibrium, as well as observations of selection coefficients as predictors of local adaptation for equilibrium and nonequilibrium systems.


Sources and sinks

The 2 Island source-sink gallery has some simulations and preliminary results concerning affects of sources and sinks on local adaptation and stability of the polymorphic equilibrium. Single island sink has some analytic results for the stability of equilibria when reciving fixed allele frequencies from a source.